See also Bibiana’s Google Scholar profile    ORCID: 0020-4435-6914

Names in bold denote group members; underlined names correspond to supervised MSc students; asterisks (*) indicate undergraduate students and mentees


33. Rönkä, K., Valkonen, J., Nokelainen, O., Rojas, B., Gordon, S., Burdfield-Steel, E. & Mappes, J. Geographic mosaic of selection by avian predators on hindwing warning colour of a polymorphic aposematic moth.  Ecology Letters.DOI: 10.1111/ele.13597 PDF

32. Fouilloux, C,. Garcia, G. & Rojas, B. Visible implant elastomer (VIE) persistence across development in a small anuran species. PeerJ 8:e9630 DOI: 10.7717/peerj.9630

31. Schulte, L. M., Ringler, E.‡, Rojas, B.‡ & Stynoski, J. L.‡ 2020. Developments in amphibian parental care research: history, present advances and future perspectives. (‡Equal contribution; listed in alphabetical order). Herpetological Monographs 34:71-97. DOI: 10.1655/HERPMONOGRAPHS-D-19-00002.1 PDF

30. Carvajal-Castro, J. D.*, López-Aguirre, Y., Ospina, A. M., Santos, J. C., Rojas, B. & Vargas-Salinas, F. 2020 Much more than a clasp: evolutionary patterns of amplexus diversity in anurans. Biological Journal of the Linnean Society 129:652-663. DOI: 10.1093/biolinnean/blaa009


29. Fouilloux, C., Ringler, E. & Rojas, B. 2019. Cannibalism. Quick Guide. Current Biology 29:R1295-R1297. DOI: 10.1016/j.cub.2019.09.068

28. Rojas, B. & Pašukonis, A. 2019. From habitat use to social behavior: natural history of a voiceless poison frog, Dendrobates tinctorius. PeerJ 7:e7648. DOI: 10.7717/peerj.7648 PDF

27. Lawrence, J. P.‡, Rojas, B.‡, Fouquet, A., Mappes, J., Blanchette , A., Saporito, R., Bosque, R. J., Courtois, E., & Noonan, B. P. 2019. Weak warning signals can persist in the absence of gene flow. (‡Equal contribution). PNAS 11619037-19045. DOI:10.1073/pnas.1901872116. PDF

Featured in: The New York Times, Science Daly,, Dr.Stevil’s BlogCosmos Magazine, ELEMENT (Russia)GreenReport (Italy).


26. Pašukonis, A., Loretto, M. C. & Rojas, B. 2019. How far do tadpoles travel in the rainforest? Parent-assisted dispersal in poison frogs. Evolutionary Ecology. DOI:10.1007/s10682-019-09994-z PDF

Featured in: Scientific American, GreenReport (Italy).


25. Rojas, B., Mappes, J. & Burdfield-Steel, E. 2019. Multiple modalities in insect warning displays have additive effects against wild avian predators. Behavioral Ecology and Sociobiology 73:37. DOI: 10.1007/s00265-019-2643-6 PDF

24. Bernal, X. E., Rojas, B., Pinto-E, M. A., Mendoza-Henao, Á. M., Herrera-Montes, A., Herrera-Montes, M. I. and Cáceres Franco, A. P. 2019. Empowering Latina scientists. Science 363:825. PDF


23. Burdfield-Steel, E., Brain, M.Rojas, B. & Mappes, J. 2018. The price of safety: food deprivation in early life influences the efficacy of chemical defence in an aposematic moth. Oikos 128:245-253. DOI:10.1111/oik.05420

22. Rojas, B., Burdfield-Steel, E., Gordon, S. P., De Pasqual, C., Hernández, L., Mappes, J., Nokelainen, O., Rönkä, K., Lindstedt, C. 2018. Multimodal aposematic signals and their emerging role in mate attraction. Frontiers in Ecology and Evolution 9:93DOI: 10.3389/fevo.2018.00093/abstract

21. Rönkä, K., Mappes, J., Kiviö, R., Salokannas, J., Michalis, C. & Rojas, B. 2018. Can multiple-model mimicry explain warning signal polymorphism in the wood tiger moth, Arctia plantaginis (Lepidoptera: Erebidae)? Biological Journal of the Linnean Society 124: 237-260. DOI: 10.1093/biolinnean/bly042 PDF

20. Burdfield-Steel, E., Pakkanen, H., Rojas, B., Galarza, J. A. &  Mappes, J. 2018. De novo synthesis of chemical defences in an aposematic moth. Journal of Insect Science 18(2):28. PDF

19. Henze, M. J., Lind, O., Mappes, J., Rojas, B. and Kelber, A. 2018. An aposematic colour‐polymorphic moth seen through the eyes of conspecifics and predators ‐ sensitivity and colour discrimination in a tiger moth. Functional Ecology 32: 1797-1809. DOI:10.1111/1365-2435.13100 PDF

18. Rönkä, K., De Pasqual, C., Mappes, J., Gordon, S. P. & Rojas, B.  2018. Colour alone matters: no predator generalisation among morphs of an aposematic moth. Animal Behaviour 135: 153–163DOI: 10.1016/j.anbehav.2017.11.015 PDF


17. Rojas, B.‡, Burdfield-Steel, E.‡, Pakkanen, H., Suisto, K., Maczka, M., Schulz, S. & Mappes, J. 2017. How to fight multiple enemies: target-specific chemical defences in an aposematic moth. (‡Equal contribution). Proceedings of the Royal Society B 284: 20171424. DOI: 10.1098/rspb.2017.1424 PDF

Featured in:  Inside ScienceDiscoverMagazineScientific American’s “60 Second Science” podcast, Phys.orgScience Daily.


16. White, T. E., Rojas, B., Mappes, J., Rautiala, P. & Kemp, D. J. 2017. Colour and luminance contrasts predict the human detection of natural stimuli in complex visual environments. Biology Letters 13: 20170375. DOI: 10.1098/rsbl.2017.0375 PDF


15. Rojas, B. 2017. Behavioural, ecological, and evolutionary aspects of diversity in frog colour patterns. Biological Reviews 92:1059-1080. DOI: 10.1111/brv.12269 PDF


14. Stynoski, J. L.‡, Schulte, L. M.‡ & Rojas, B.‡ 2015. Poison frogs. Quick Guide. Current Biology 25:R1026–R1028.(‡ Equal contribution) PDF

13. Rojas, B. 2015. Mind the gap: treefalls as drivers of parental tradeoffs. Ecology & Evolution. DOI: 10.1002/ece3.1648 PDF

Featured in: Sciences et Avenir (France)


12. Rojas, B.‡, Gordon, S. P.‡ & Mappes, J. 2015. Frequency-dependent flight activity in the colour polymorphic wood tiger moth. Current Zoology 61:765-772. Special issue on ‘Anti-predator coloration and behavior’ (‡ Equal contribution) PDF

11. Gordon, S. P., Kokko, H., Rojas, B., Nokelainen, O. & Mappes, J. 2015. Colour polymorphism torn apart by opposing positive frequency-dependent selection, yet maintained in space. Journal of Animal Ecology. DOI: 10.1111/1365-2656.12416 PDF

Featured in a Special Virtual Issue on Evolutionary Ecology (editor’s choice) in the Journal of Animal Ecology

10. Exnerová, A., Jezová, D., Štys, P., Doktorovová, L., Rojas, B. & Mappes, J. 2015. Different reactions to aposematic prey in 2 geographically distant populations of great tits. Behavioral Ecology 26:1361-1370. DOI: 10.1093/beheco/arv086 PDF

9. Hämäläinen, L., Valkonen, J., Mappes, J. & Rojas, B. 2015. Visual illusions in predator-prey interactions: birds find moving patterned prey harder to catch. Animal Cognition 18:1059-1068. DOI: 10.1007/s10071-015-0874-0 PDF

8. Rojas, B., Valkonen, J. & Nokelainen, O. 2015. Aposematism. Quick Guide. Current Biology 25:R350-R351. PDF


7. Rojas, B., Rautiala, P. & Mappes, J. 2014. Differential detectability of polymorphic warning signals under varying light environments. Behavioural Processes 109(B): 164-172. Special issue on ‘Animal Cognition in the Wild’. PDF


6. Rojas, B., Devillechabrolle, J. & Endler, J. A. 2014. Paradox lost: colour pattern and movement are associated in an aposematic frog. Biology Letters 10: 20140193. PDF

Featured in:, Science News, Daily Mail (UK), Sydney Morning Herald (Australia), Nature World News, Spektrum (Germany)France Inter.

5. Rojas, B. 2014. Strange parental decisions: fathers of the dyeing poison frog deposit their tadpoles in pools occupied by large cannibals. Behavioral Ecology and Sociobiology 68:551-559. PDF

Featured in: Science Magazine, Science Daily,, Springer Select, The Scientist magazine, Der Standard (Austria), Scinexx (Germany).



4. Rojas, B. & Endler, J. A. 2013. Sexual dimorphism and intra-populational colour pattern variation in the aposematic frog Dendrobates tinctorius. Special Issue on the ‘Evolutionary Ecology of Poison Frogs’. Evolutionary Ecology 27:739-753. PDF

Before 2013

3. Ringler, E., Rojas, B., Ringler, M. & Hödl, W. 2012. Characterisation of nine polymorphic microsatellite loci in the dyeing poison frog Dendrobates tinctorius (Dendrobatidae), and their cross-species utility in two other dendrobatoid species. Herpetological Journal 22: 265-267. PDF

2. Endler, J. A. & Rojas, B. 2009. The spatial pattern of natural selection when selection depends on experience. American Naturalist 173: E62-E78. PDF

1. Rojas, B., Amézquita, A. & Delgadillo, A. 2006. Matching and symmetry in the frequency recognition curve of the poison frog Epipedobates trivittatus. Ethology 112: 564-571. PDF


Fouilloux, C., Fromhage, L., Valkonen, J .& Rojas, B. Size-dependent tradeoffs in cannibalistic behaviour towards kin. Under review.

In preparation 

Rojas, B., Valkonen, J. Ringler, M. & Endler, J. A. Bold invaders are winners: poison frogs with simple aposematic colour patterns lead tree-fall gap invasions and have higher survival.

Carvajal-Castro, J. D.*, Vargas-Salinas, F., Casas-Cardona, S., Rojas, B. & Santos, J.C. The evolution of parental care in poison frogs.

Rojas, B., Rönkä, K., Nokelainen, O., Valkonen, J. & Mappes, J. Red or dead: imperfect Müllerian mimicry between burnet and red, not yellow, wood tiger moths.

Ottocento, C., Rojas, B., Burdfield-Steel, E., Furlanetto, M.Nokelainen, O. & Mappes, J. Strategic resource allocation in male wood tiger moth Arctia plantaginis: the role of early-life resource availability in warning signal and chemical defence.

Other publications


Fouilloux, C., Garcia, G. & Rojas, B. Visible implant elastomer (VIE) success in early larval stages of a tropical amphibian species. bioRxiv 2020.04.29.057232; DOI:10.1101/2020.04.29.057232

Rönkä, K., Valkonen, J., Nokelainen, O., Rojas, B., Gordon, S., Burdfield-Steel, E. & Mappes, J. 2020. Geographic mosaic of selection by avian predators on hindwing warning colour of a polymorphic aposematic moth.

Carvajal-Castro, J. D.*, López-Aguirre, Y., Ospina, A. M., Santos, J. C., Rojas, B. & Vargas-Salinas, F. 2020. Much more than a clasp:  evolutionary pattern of amplexus diversity in anurans.

Rojas, B. & Pašukonis, A. 2019. From habitat use to social behavior: natural history of a voiceless poison frog, Dendrobates tinctorius. bioRxiv 


5. Rojas, B. 2018. Comentario científico: Hacia un estudio integral del papel de los patrones de coloración en las interacciones entre depredadores y presas. Boletín Colombiano de Biología Evolutiva 6: 14-15. PDF

4. Rojas, B. & Burdfield-Steel, E. Predator Defense. 2017. In: J. Vonk & T. K. Shackelford (Eds.). Encyclopaedia of Animal Cognition and Behavior. Springer International Publishing.  DOI: 10.1007/978-3-319-47829-6_708-1 PDF

3. Nokelainen, O., Rojas, B., & Valkonen, J. Camouflage. 2017. In: T. K. Shackelford & V. A. Weekes-Shackelford (Eds.). Encyclopaedia of Evolutionary Psychological Science. Springer International PublishingDOI:10.1007/978-3-319-16999-6_2665-1  PDF

2. Rojas, B., Nokelainen, O., & Valkonen, J. Aposematism. 2017. In: T. K. Shackelford & V. A. Weekes-Shackelford (Eds.). Encyclopaedia of Evolutionary Psychological Science. Springer International Publishing. DOI:10.1007/978-3-319-16999-6_2669-1  PDF

1. Burdfield-Steel, E. & Rojas, B. 2017. Doubling down. BIOSPHERE Magazine 29:29-36. PDF